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Table 2 Communities based on the cluster analysis (labelled with roman numerals), location features (exposure of the shore; vertical zone) and communities based on the inverse analysis; these communities were named after characteristic and dominant species restricted to a specific community (short names of the communities are in bold)

From: Community composition of the rocky intertidal at Helgoland (German Bight, North Sea)

Communities based on cluster analysis

Location features

Communities based on inverse analysis

Biotope name according to Bartsch and Tittley (2004)

Exposure

Zone

Community I

Semi-exposed

High intertidal

Ulva spp.

Enteromorpha spp. on freshwater-influenced and/or unstable upper eulittoral rock

Community II

Semi-exposed

Mid intertidal

Mytilusedulis–Littorinalittorea–Ralfsia verrucosa

Mytilus edulis and Fucusvesiculosus on moderately exposed mid eulittoral rock

Community III

Semi-exposed to exposed

Mid to low intertidal

Corallina officinalis– Fucus serratus

Corallina officinalis, coralline crusts and brown seaweeds in shallow eulittoral rockpools

Dense Fucus serratus on moderately exposed to very sheltered full salinity lower eulittoral rock

Fucus serratus and red seaweeds on moderately exposed lower eulittoral rock

Mastocarpus stellatus and Chondrus crispus on very exposed to moderately exposed lower eulittoral rock

Community IV

Sheltered

Mid to low intertidal (horizontal)

Cryptosula pallasiana–Dynamena pumila–Ascophyllumnodosum

Ascophyllum nodosum on very sheltered mid eulittoral rock

Community V

Semi-exposed to exposed

Low intertidal

Rhodothamniellafloridula–tube-building polychaetes–Ulva lactuca

Rhodothamniellafloridula on sand-scoured lower eulittoral rock

Community VI

Sheltered

High intertidal (vertical)

Balanus spp.–Fucus spiralis–Porphyra spp.

Semibalanus balanoides, Patella vulgata and Littorina spp. on exposed to moderately exposed or vertical sheltered eulittoral rock

  1. The last column shows the biotope name according to Bartsch and Tittley (2004)