Table 1 Abundance (mean percent cover and SE) of the species found at Nord-Ost Hafen at elevation zones 1 (between −10 cm and 20 cm of elevation, relative to chart datum), 2 (20–50 cm), 3 (50–80 cm), 4 (80–110 cm), 5 (110–140 cm), 6 (140–170 cm), 7 (170–200 cm), 8 (200–230 cm), and 9 (230–260 cm)
| Â | Elevation zone 1 | Elevation zone 2 | Elevation zone 3 | Elevation zone 4 | Elevation zone 5 | |||||
|---|---|---|---|---|---|---|---|---|---|---|
Mean | SE | Mean | SE | Mean | SE | Mean | SE | Mean | SE | |
Algae | ||||||||||
Ascophyllum nodosum* | 0.25 | 0.25 | – | – | 2.81 | 2.81 | – | – | – | – |
Blidingia sp. | – | – | – | – | – | – | – | – | – | – |
Ceramium rubrum | 10.79 | 3.71 | 2.13 | 1.00 | 0.69 | 0.57 | 0.62 | 0.45 | 0.08 | 0.05 |
Chaetomorpha sp. | 0.75 | 0.22 | 1.27 | 0.98 | 0.19 | 0.07 | 0.65 | 0.25 | 0.48 | 0.16 |
Chondrus crispus | 18.25 | 2.28 | 15.13 | 2.33 | 6.04 | 2.42 | 3.58 | 1.21 | 0.65 | 0.31 |
Cladophora rupestris | 4.67 | 2.43 | 6.31 | 2.48 | 2.00 | 0.48 | 2.67 | 0.86 | 1.04 | 0.44 |
Cladostephus spongiosus* | 0.08 | 0.08 | – | – | – | – | – | – | – | – |
Cystoclonium purpureum* | 0.58 | 0.37 | 0.15 | 0.08 | – | – | – | – | – | – |
Desmarestia aculeata* | 0.13 | 0.09 | 0.04 | 0.04 | – | – | – | – | – | – |
Elachista fucicola | 0.60 | 0.37 | 4.38 | 1.19 | 2.35 | 1.44 | 0.73 | 0.73 | 0.15 | 0.12 |
Encrusting green algae | 0.08 | 0.08 | 0.29 | 0.25 | 0.67 | 0.31 | 4.33 | 2.26 | 4.46 | 1.38 |
Filamentous green algae | 1.58 | 0.75 | 1.63 | 0.81 | 1.27 | 0.56 | 0.54 | 0.26 | 0.65 | 0.39 |
Filamentous brown algae | 0.60 | 0.29 | – | – | – | – | 2.85 | 2.76 | 0.12 | 0.12 |
Fucus serratus | 38.27 | 10.87 | 62.75 | 6.59 | 66.77 | 6.79 | 45.54 | 10.07 | 34.58 | 8.77 |
Fucus spiralis | – | – | – | – | – | – | – | – | – | – |
Fucus vesiculosus | – | – | – | – | 4.27 | 2.67 | 1.46 | 1.38 | 6.15 | 3.76 |
Giffordia sp. | 1.50 | 0.96 | 11.00 | 2.39 | 8.90 | 3.64 | 2.71 | 1.24 | 0.37 | 0.17 |
Hildenbrandia rubra | – | – | 0.17 | 0.07 | 0.14 | 0.07 | 0.04 | 0.04 | 0.04 | 0.04 |
Laminaria digitata* | 38.54 | 11.45 | 0.63 | 0.63 | 2.12 | 1.59 | – | – | 0.08 | 0.05 |
Laminaria hyperborea* | 19.81 | 9.51 | 3.13 | 1.16 | 0.46 | 0.46 | – | – | – | – |
Mastocarpus stellatus | 5.96 | 1.53 | 19.38 | 3.25 | 38.54 | 6.60 | 60.92 | 8.10 | 76.46 | 9.59 |
Membranoptera alata* | 2.54 | 0.86 | 0.31 | 0.12 | 0.19 | 0.07 | 0.04 | 0.04 | – | – |
Petalonia fascia | – | – | – | – | 0.23 | 0.23 | – | – | – | – |
Phymatolithon sp. | 3.98 | 1.58 | 4.63 | 0.99 | 11.46 | 4.09 | 4.65 | 1.67 | 4.89 | 2.14 |
Plocamium cartilagineum* | 0.04 | 0.04 | – | – | – | – | – | – | – | – |
Plumaria elegans* | 0.17 | 0.11 | 0.08 | 0.08 | – | – | – | – | – | – |
Polysiphonia sp.* | 1.21 | 0.99 | 0.06 | 0.06 | 0.04 | 0.04 | – | – | – | – |
Porphyra sp. | 0.71 | 0.26 | 0.79 | 0.28 | 0.46 | 0.34 | 0.15 | 0.12 | – | – |
Ralfsia sp. | – | – | – | – | – | – | – | – | 0.96 | 0.92 |
Rhodomela confervoides | 0.23 | 0.12 | – | – | – | – | – | – | – | – |
Rhodothamniella floridula | 24.75 | 8.40 | 15.67 | 5.16 | 14.27 | 7.65 | 5.85 | 4.06 | 0.96 | 0.52 |
Saccharina latissima | 14.50 | 8.66 | – | – | – | – | 0.62 | 0.49 | – | – |
Sargassum muticum* | 0.08 | 0.08 | – | – | – | – | – | – | – | – |
Spongonema tomentosum | – | – | 0.42 | 0.34 | – | – | 0.46 | 0.46 | 0.08 | 0.08 |
Ulva sp. 1 | 6.63 | 2.41 | 7.96 | 4.38 | 4.69 | 1.50 | 7.08 | 4.56 | 0.42 | 0.13 |
Ulva sp. 2 | – | – | – | – | 0.08 | 0.08 | 0.23 | 0.23 | – | – |
Invertebrates | ||||||||||
Alcyonidium polyoum | – | – | 0.08 | 0.08 | 0.27 | 0.23 | 0.04 | 0.04 | – | – |
Amphipoda | 0.23 | 0.08 | 0.38 | 0.07 | 0.42 | 0.05 | 0.52 | 0.02 | 0.54 | 0.03 |
Annelida | – | – | – | – | – | – | 0.04 | 0.04 | – | – |
Balanus balanus | 1.75 | 1.66 | 0.43 | 0.31 | 0.42 | 0.35 | 10.03 | 6.79 | 19.02 | 7.54 |
Balanus crenatus | – | – | – | – | 0.04 | 0.04 | – | – | 0.23 | 0.23 |
Balanus improvisus | 2.54 | 1.16 | 2.39 | 0.66 | 18.36 | 7.96 | 12.01 | 6.04 | 12.16 | 6.43 |
Carcinus maenas | – | – | 0.04 | 0.04 | – | – | – | – | 0.12 | 0.12 |
Clava multicornis | – | – | 0.13 | 0.07 | 0.04 | 0.04 | 0.04 | 0.04 | 0.04 | 0.04 |
Crassostrea gigas | 0.04 | 0.04 | 0.83 | 0.06 | 0.33 | 0.16 | 1.80 | 0.26 | 0.40 | 0.24 |
Dipteran larvae | – | – | – | – | – | – | 0.04 | 0.04 | – | – |
Dynamena pumila | 0.25 | 0.10 | 0.44 | 0.12 | 0.85 | 0.39 | 2.14 | 1.05 | 2.12 | 0.87 |
Electra pilosa* | 4.67 | 1.38 | 0.90 | 0.25 | 0.19 | 0.07 | 0.08 | 0.08 | <0.01 | <0.01 |
Elminius modestus | 0.83 | 0.83 | 2.28 | 1.26 | 8.97 | 3.54 | 25.43 | 5.99 | 25.20 | 7.00 |
Flustrella hispida | 8.13 | 1.56 | 11.48 | 1.23 | 7.62 | 1.75 | 12.00 | 3.12 | 11.08 | 3.02 |
Gibbula cineraria* | 0.08 | 0.06 | 0.08 | 0.06 | – | – | – | – | – | – |
Halichondria panicea* | 0.04 | 0.04 | – | – | – | – | – | – | – | – |
Hydrobia sp. | 0.04 | 0.04 | – | – | 0.04 | 0.04 | – | – | – | – |
Idotea granulosa | 0.17 | 0.07 | 0.40 | 0.07 | 0.23 | 0.07 | 0.15 | 0.07 | 0.12 | 0.06 |
Janua pagenstecheri | 0.04 | 0.04 | 0.77 | 0.35 | 1.40 | 0.30 | 2.39 | 0.64 | 1.90 | 0.72 |
Lacuna vincta | 0.21 | 0.09 | – | – | – | – | – | – | – | – |
Laumeda flexulosa | – | – | – | – | – | – | – | – | 0.04 | 0.04 |
Leucosolenia sp.* | 0.19 | 0.10 | – | – | 0.06 | 0.06 | 0.40 | 0.19 | 0.08 | 0.08 |
Littorina littorea | – | – | 0.13 | 0.08 | 0.04 | 0.04 | 0.10 | 0.07 | 0.15 | 0.08 |
Littorina obtusata | 0.29 | 0.07 | 0.69 | 0.07 | 0.71 | 0.20 | 0.67 | 0.10 | 0.54 | 0.06 |
Littorina saxatilis | – | – | – | – | 0.08 | 0.05 | 0.12 | 0.06 | 0.12 | 0.06 |
Membranipora membranacea* | 0.04 | 0.04 | – | – | – | – | – | – | – | – |
Mytilus edulis | 0.08 | 0.06 | 0.13 | 0.07 | 0.12 | 0.06 | 0.19 | 0.12 | 0.25 | 0.10 |
Obelia geniculata | – | – | – | – | – | – | – | – | 0.04 | 0.04 |
Pomatoceros triqueter | – | – | – | – | 0.04 | 0.04 | 0.04 | 0.04 | 0.04 | 0.04 |
Semibalanus balanoides | – | – | – | – | – | – | – | – | 0.04 | 0.04 |
Spirorbis spirorbis* | 0.15 | 0.08 | 0.52 | 0.28 | 0.69 | 0.45 | 0.23 | 0.16 | – | – |
Sycon ciliatum* | 0.06 | 0.06 | – | – | – | – | – | – | – | – |
Verruca stroemia | 19.71 | 8.79 | 10.32 | 3.56 | 11.42 | 4.28 | 19.07 | 9.22 | 3.16 | 1.80 |
Relevant group information | ||||||||||
All species | 236.83 | 12.37 | 189.86 | 11.95 | 220.10 | 16.00 | 232.06 | 13.59 | 209.99 | 7.95 |
Fucus spp. | 38.27 | 10.87 | 62.75 | 6.59 | 71.04 | 7.99 | 47.00 | 10.40 | 40.73 | 11.01 |
Fucus spp., Mastocarpus stellatus, and Chondrus crispus | 62.48 | 11.74 | 97.25 | 5.70 | 115.62 | 9.57 | 111.50 | 6.03 | 117.85 | 6.93 |
Littorina spp. | 0.29 | 0.07 | 0.81 | 0.12 | 0.83 | 0.20 | 0.89 | 0.17 | 0.81 | 0.12 |
| Â | Elevation zone 6 | Elevation zone 7 | Elevation zone 8 | Elevation zone 9 | ||||
|---|---|---|---|---|---|---|---|---|
Mean | SE | Mean | SE | Mean | SE | Mean | SE | |
Algae | ||||||||
Ascophyllum nodosum* | – | – | – | – | – | – | – | – |
Blidingia sp. | – | – | 7.73 | 3.74 | 0.08 | 0.08 | – | – |
Ceramium rubrum | – | – | – | – | – | – | – | – |
Chaetomorpha sp. | 0.31 | 0.09 | 0.04 | 0.04 | – | – | – | – |
Chondrus crispus | 0.17 | 0.10 | – | – | 0.35 | 0.35 | – | – |
Cladophora rupestris | 0.35 | 0.24 | 0.12 | 0.08 | – | – | – | – |
Cladostephus spongiosus* | – | – | – | – | – | – | – | – |
Cystoclonium purpureum* | – | – | – | – | – | – | – | – |
Desmarestia aculeata* | – | – | – | – | – | – | – | – |
Elachista fucicola | – | – | – | – | – | – | – | – |
Encrusting green algae | 15.54 | 3.75 | 38.50 | 9.36 | 14.15 | 7.40 | 3.39 | 2.33 |
Filamentous green algae | 0.58 | 0.58 | 2.15 | 1.92 | 12.15 | 7.81 | – | – |
Filamentous brown algae | – | – | – | – | – | – | – | – |
Fucus serratus | 10.00 | 3.04 | 4.04 | 2.84 | – | – | – | – |
Fucus spiralis | 0.46 | 0.46 | 0.39 | 0.27 | – | – | – | – |
Fucus vesiculosus | 25.77 | 7.28 | 19.00 | 6.47 | 0.85 | 0.68 | – | – |
Giffordia sp. | – | – | – | – | – | – | – | – |
Hildenbrandia rubra | 0.65 | 0.23 | 1.00 | 0.24 | 0.27 | 0.23 | – | – |
Laminaria digitata* | – | – | – | – | – | – | – | – |
Laminaria hyperborea* | – | – | – | – | – | – | – | – |
Mastocarpus stellatus | 77.08 | 5.14 | 37.62 | 9.74 | 1.19 | 1.15 | – | – |
Membranoptera alata* | – | – | – | – | – | – | – | – |
Petalonia fascia | – | – | – | – | – | – | – | – |
Phymatolithon sp. | 2.65 | 1.06 | 1.15 | 0.70 | – | – | – | – |
Plocamium cartilagineum* | – | – | – | – | – | – | – | – |
Plumaria elegans* | – | – | – | – | – | – | – | – |
Polysiphonia sp.* | – | – | – | – | – | – | – | – |
Porphyra sp. | – | – | 0.46 | 0.23 | 0.08 | 0.05 | – | – |
Ralfsia sp. | 2.89 | 1.27 | 43.50 | 10.49 | 87.08 | 3.12 | 78.62 | 4.85 |
Rhodomela confervoides | – | – | – | – | – | – | – | – |
Rhodothamniella floridula | – | – | 0.58 | 0.58 | – | – | – | – |
Saccharina latissima | – | – | – | – | – | – | – | – |
Sargassum muticum* | – | – | – | – | – | – | – | – |
Spongonema tomentosum | – | – | – | – | – | – | – | – |
Ulva sp. 1 | 0.04 | 0.04 | – | – | – | – | – | – |
Ulva sp. 2 | – | – | – | – | – | – | – | – |
Invertebrates | ||||||||
Alcyonidium polyoum | – | – | – | – | – | – | – | – |
Amphipoda | 0.73 | 0.12 | 0.58 | 0.11 | 0.12 | 0.06 | – | – |
Annelida | – | – | – | – | – | – | – | – |
Balanus balanus | 5.12 | 2.96 | 3.12 | 1.99 | 0.04 | 0.04 | – | – |
Balanus crenatus | 0.28 | 0.14 | 2.91 | 1.30 | 1.54 | 0.55 | 0.35 | 0.23 |
Balanus improvisus | 4.79 | 3.18 | 0.89 | 0.46 | 0.04 | 0.04 | – | – |
Carcinus maenas | – | – | – | – | – | – | – | – |
Clava multicornis | 0.08 | 0.05 | – | – | – | – | – | – |
Crassostrea gigas | 1.29 | 0.37 | 0.65 | 0.58 | – | – | – | – |
Dipteran larvae | – | – | 0.04 | 0.04 | – | – | – | – |
Dynamena pumila | 0.50 | 0.12 | 0.08 | 0.05 | – | – | – | – |
Electra pilosa* | 0.04 | 0.04 | – | – | – | – | – | – |
Elminius modestus | 15.50 | 6.56 | 14.31 | 4.88 | 2.19 | 0.91 | 0.77 | 0.69 |
Flustrella hispida | 14.87 | 5.53 | 1.27 | 0.76 | – | – | – | – |
Gibbula cineraria* | – | – | – | – | – | – | – | – |
Halichondria panicea* | – | – | – | – | – | – | – | – |
Hydrobia sp. | – | – | – | – | – | – | – | – |
Idotea granulosa | 0.12 | 0.06 | 0.04 | 0.04 | – | – | – | – |
Janua pagenstecheri | 3.50 | 0.95 | 0.19 | 0.11 | – | – | – | – |
Lacuna vincta | – | – | – | – | – | – | – | – |
Laumeda flexulosa | – | – | – | – | – | – | – | – |
Leucosolenia sp.* | 0.04 | 0.04 | – | – | – | – | – | – |
Littorina littorea | 0.06 | 0.06 | 0.15 | 0.08 | 0.17 | 0.10 | 0.17 | 0.13 |
Littorina obtusata | 0.25 | 0.25 | 0.46 | 0.08 | 0.04 | 0.04 | – | – |
Littorina saxatilis | 0.15 | 0.07 | 0.33 | 0.10 | 0.37 | 0.07 | 0.19 | 0.07 |
Membranipora membranacea* | – | – | – | – | – | – | – | – |
Mytilus edulis | 0.27 | 0.07 | 0.06 | 0.06 | – | – | – | – |
Obelia geniculata | 0.04 | 0.04 | 0.04 | 0.04 | – | – | – | – |
Pomatoceros triqueter | – | – | – | – | – | – | – | – |
Semibalanus balanoides | 0.36 | 0.17 | 2.44 | 1.89 | 0.19 | 0.09 | 0.04 | 0.04 |
Spirorbis spirorbis* | – | – | – | – | – | – | – | – |
Sycon ciliatum* | – | – | – | – | – | – | – | – |
Verruca stroemia | – | – | – | – | – | – | – | – |
Relevant group information | ||||||||
All species | 184.92 | 11.50 | 183.81 | 12.74 | 120.89 | 11.32 | 83.15 | 4.70 |
Fucus spp. | 36.23 | 7.61 | 23.42 | 7.91 | 0.85 | 0.68 | – | – |
Fucus spp., Mastocarpus stellatus, and Chondrus crispus | 113.48 | 7.34 | 61.04 | 16.24 | 2.39 | 1.83 | – | – |
Littorina spp. | 0.94 | 0.19 | 0.94 | 0.20 | 0.58 | 0.15 | 0.37 | 0.16 |